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| Family: Liocranidae (Spiny-legged Sac Spiders) |
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| Genus: Agraecina Simon, 1932 |
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| Agraecina canariensis Wunderlich, 1992 |
| Range: Spain (Canary Islands) (van Helsdingen 2009.1). Global range: Canary Is (Platnick 10.0). |
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| Agraecina cristiani (Georgescu, 1989) |
| Range: Romania (van Helsdingen 2009.1). Global range: Romania (Platnick 10.0). |
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| Agraecina hodna Bosmans, 1999 |
| Range: Macedonia? (van Helsdingen 2009.1). Global range: Algeria, Macedonia (Platnick 10.0). |
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| Agraecina lineata (Simon, 1878) |
| Range: Belgium, France (Corsica), France (Mainland), Italy (Mainland), Portugal (Mainland), Russia (Southern European), Spain (Mainland) (van Helsdingen 2009.1). Global range: Western Mediterranean (Platnick 10.0). |
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| Genus: Agroeca Westring, 1861 |
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| Characters of genus: The carapace usually with clear markings in the shape of characteristic angular loops radiating from the fovea. The abdomen with vague v- or w-shaped pattern or dark oblique spots or bars arranged in rows. The posterior row of eyes markedly procurved. Metatarsus I and II with three pairs of ventral spines. The Danish species can be identified in the field by their bodycolour and size. |
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| Agroeca annulipes Simon, 1878 |
| Range: France (Corsica), Italy (Sardinia), Spain (Mainland) (van Helsdingen 2009.1). Global range: Spain, Corsica, Sardinia, Morocco, Algeria (Platnick 10.0). |
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| Agroeca brunnea (Blackwall, 1833) |
| Description: The largest species of the genus. Carapace reddish-brown with golden hairs, and dark markings. Abdomen golden reddish-brown somewhat shiny. Similar to A. proxima which overlap in habitat but is smaller. The eggsacks are very characteristic and look like small inverted wineglasses placed in the vegetation, and are encountered more frequently than the spider itself. The eggsacks may be snowy-white or camouflaged with earth, thereby much less conspiceous. Size: Female 7-8 mm; male 6-7 mm. Maturity: All year. Habitat: Damp or dry places with low vegetation, usually near or within forests, sometimes also moors and grassland. Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Hungary, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0). |
Juvenile. |
Juvenile. |
Juvenile. |
Female. |
Female. |
Female abdomen. |
Female. |
Female. |
Male. |
Male. |
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| Agroeca cuprea Menge, 1873 |
| Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Liechtenstein, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Central Asia (Platnick 10.0). |
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| Agroeca dentigera Kulczynski, 1913 |
| Range: Belarus, Belgium, Finland, Germany, Great Britain (Mainland), Lithuania, Netherlands, Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Southern European), Sweden, Ukraine (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0). |
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| Agroeca gaunitzi Tullgren, 1952 |
| Range: Sweden (van Helsdingen 2009.1). Global range: Sweden (Platnick 10.0). |
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| Agroeca inopina O. P.-Cambridge, 1886 |
| Range: Belgium, Bulgaria, France (Corsica), France (Mainland), Great Britain (Channel Islands), Great Britain (Mainland), Italy (Mainland), Netherlands, Portugal (Mainland), Slovakia, Spain (Mainland), Switzerland (van Helsdingen 2009.1). Global range: Europe, Algeria (Platnick 10.0). |
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| Agroeca lusatica (L. Koch, 1875) |
| Description: Agroeca lusatica is slightly larger than its close relative Agroeca proxima, which is found in the same habitat however usually slightly further inland. Size: Female 6-7 mm; male 5-6 mm. Maturity: Autumn. Habitat: In Denmark this species is only known from coastal dunes. In Europe it is also associated with woodland. Range: Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Hungary, Italy (Mainland), Lithuania, Macedonia, Netherlands, Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (NW. European), Russia (Southern European), Slovakia, Spain (Mainland), Sweden, Ukraine (van Helsdingen 2009.1). Global range: Europe to Kazakhstan (Platnick 10.0). |
Female. |
Male. |
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| Agroeca maculata L. Koch, 1879 |
| Range: Russia (Central European), Russia (Eastern European) (van Helsdingen 2009.1). Global range: Russia, Kazakhstan (Platnick 10.0). |
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| Agroeca makarovae Esyunin, 2008 |
| Range: Russia (Eastern European) (van Helsdingen 2009.1). Global range: Russia (Platnick 10.0). |
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| Agroeca proxima (O. P.-Cambridge, 1871) |
| Description: Rather similar to A. brunnea but smaller. The eggsack is much narrower than that of A. brunnea. Size: Female 5.5-7.5 mm; male 4-5.5 mm. Habitat: Primarily heather and coastal dunes, but sometimes also in the same habitat as A. brunnea. Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Hungary, Ireland, Italy (Mainland), Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0). |
Male. |
Female. |
Female. |
Male. |
Male. |
Male abdominal markings. |
Male. |
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| Genus: Apostenus Westring, 1851 |
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| Apostenus annulipedes Wunderlich, 1987 |
| Range: Spain (Canary Islands) (van Helsdingen 2009.1). Global range: Canary Is (Platnick 10.0). |
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| Apostenus fuscus Westring, 1851 |
| Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Finland, France (Mainland), Germany, Great Britain (Mainland), Hungary, Italy (Mainland), Liechtenstein, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0). |
Female. |
Female. |
Male. |
Male. |
Male. |
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| Apostenus gomerensis Wunderlich, 1992 |
| Range: Spain (Canary Islands) (van Helsdingen 2009.1). Global range: Canary Is (Platnick 10.0). |
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| Apostenus grancanariensis Wunderlich, 1992 |
| Range: Spain (Canary Islands) (van Helsdingen 2009.1). Global range: Canary Is (Platnick 10.0). |
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| Apostenus humilis Simon, 1932 |
| Range: France (Mainland), Portugal (Mainland) (van Helsdingen 2009.1). Global range: France (Platnick 10.0). |
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| Apostenus ochraceus Hadjissarantos, 1940 |
| Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0). |
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| Apostenus palmensis Wunderlich, 1992 |
| Range: Spain (Canary Islands) (van Helsdingen 2009.1). Global range: Canary Is (Platnick 10.0). |
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| Genus: Brachyanillus Simon, 1913 |
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| Brachyanillus liocraninus Simon, 1913 |
| Range: Spain (Mainland) (van Helsdingen 2009.1). Global range: Spain, Algeria (Platnick 10.0). |
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| Genus: Cybaeodes Simon, 1878 |
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| Cybaeodes avolensis Platnick & Di Franco, 1992 |
| Range: Italy (Sicily) (van Helsdingen 2009.1). Global range: Sicily (Platnick 10.0). |
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| Cybaeodes madidus Simon, 1914 |
| Range: France (Mainland) (van Helsdingen 2009.1). Global range: France (Platnick 10.0). |
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| Cybaeodes mallorcensis Wunderlich, 2008 |
| Range: Spain (Balearic Islands) (van Helsdingen 2009.1). Global range: Mallorca (Platnick 10.0). |
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| Cybaeodes marinae Di Franco, 1989 |
| Range: Italy (Mainland), Italy (Sicily) (van Helsdingen 2009.1). Global range: Italy (Platnick 10.0). |
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| Cybaeodes molara (Roewer, 1960) |
| Range: Italy (Sicily) (van Helsdingen 2009.1). Global range: Sicily (Platnick 10.0). |
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| Cybaeodes sardus Platnick & Di Franco, 1992 |
| Range: Italy (Sardinia) (van Helsdingen 2009.1). Global range: Sardinia (Platnick 10.0). |
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| Cybaeodes testaceus Simon, 1878 |
| Range: France (Corsica) (van Helsdingen 2009.1). Global range: France, Corsica (Platnick 10.0). |
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| Genus: Liocranoeca Wunderlich, 1999 |
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| Liocranoeca spasskyi Ponomarev, 2007 |
| Range: Russia (Southern European) (van Helsdingen 2009.1). Global range: Russia (Platnick 10.0). |
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| Liocranoeca striata (Kulczynski, 1882) |
| Range: Austria, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Liechtenstein, Lithuania, Macedonia, Netherlands, Poland, Romania, Russia (Central European), Russia (Southern European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0). |
Subadult female. |
Female. |
Female. |
Subadult female. |
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| Liocranoeca striata gracilior (Kulczynski, 1898) |
| Range: Germany, Hungary, Switzerland (van Helsdingen 2009.1). Global range: Switzerland, Germany, Hungary (Platnick 10.0). |
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| Genus: Liocranum L. Koch, 1866 |
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| Characters of genus: These spiders are the largest representatives of the family in the region. The eye group is wider than in tte remaining genera and therefore the spiders resemble species of Clubionidae. |
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| Liocranum apertum Denis, 1960 |
| Range: France (Mainland) (van Helsdingen 2009.1). Global range: France (Platnick 10.0). |
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| Liocranum concolor Simon, 1878 |
| Range: France (Corsica) (van Helsdingen 2009.1). Global range: Corsica (Platnick 10.0). |
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| Liocranum giersbergi Kraus, 1955 |
| Range: Italy (Sardinia) (van Helsdingen 2009.1). Global range: Sardinia (Platnick 10.0). |
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| Liocranum kochi Herman, 1879 |
| Range: Hungary, Slovakia? (van Helsdingen 2009.1). Global range: Hungary (Platnick 10.0). |
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| Liocranum majus Simon, 1878 |
| Range: Portugal (Mainland), Spain (Balearic Islands), Spain (Mainland) (van Helsdingen 2009.1). Global range: Spain (Platnick 10.0). |
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| Liocranum ochraceum L. Koch, 1867 |
| Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Corfu (Platnick 10.0). |
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| Liocranum perarmatum Kulczynski, 1897 |
| Range: Croatia, Slovenia (van Helsdingen 2009.1). Global range: Slovenia, Croatia (Platnick 10.0). |
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| Liocranum rupicola (Walckenaer, 1830) |
| Description: Carapace is purplereddish-brown with dentate lateral bands and dark margins. The abdomen is of the same colour with a dark grey cardiac mark followed by chevrons all of which are flanked with dark spots. Legs are greyish-brown annulated with dark grey. Size: Female 6-8.5 mm; male 5.5-6 mm. Maturity: Spring and autumn. Habitat: Under stones in dry conditions. Range: Austria, Belgium, Bosnia/Herzegowina, Bulgaria, Croatia, Czech Republic, Denmark, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Hungary, Ireland, Italy (Mainland), Italy (Sardinia), Liechtenstein, Macedonia, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European)?, Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0). |
Subadult male. |
Juvenile. |
Juvenile. |
Male. |
Male. |
Male. |
Male. |
Male. |
Male abdominal markings. |
Male. |
Male. |
Male. |
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| Liocranum segmentatum Simon, 1878 |
| Range: France (Mainland), Italy (Mainland), Portugal (Mainland) (van Helsdingen 2009.1). Global range: France (Platnick 10.0). |
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| Liocranum variabilis Wunderlich, 2008 |
| Range: Spain (Balearic Islands) (van Helsdingen 2009.1). Global range: Mallorca (Platnick 10.0). |
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| Genus: Macedoniella Drensky, 1935 |
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| Macedoniella karamani Drensky, 1935 (Nomen dubium) |
| Global range: Balkans (Platnick 10.0). |
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| Genus: Mesiotelus Simon, 1897 |
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| Mesiotelus annulipes (Kulczynski, 1897) |
| Range: Bulgaria, Croatia, Hungary, Slovakia, Yugoslavia (van Helsdingen 2009.1). Global range: Hungary, Balkans, Turkey (Platnick 10.0). |
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| Mesiotelus cyprius Kulczynski, 1908 |
| Range: Bulgaria, Cyprus, Macedonia (van Helsdingen 2009.1). Global range: Cyprus (Platnick 10.0). |
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| Mesiotelus cyprius scopensis Drensky, 1935 |
| Range: Bulgaria, Greece (Mainland), Macedonia (van Helsdingen 2009.1). Global range: Greece, Bulgaria (Platnick 10.0). |
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| Mesiotelus grancanariensis Wunderlich, 1992 |
| Range: Spain (Canary Islands) (van Helsdingen 2009.1). Global range: Canary Is (Platnick 10.0). |
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| Mesiotelus maderianus Kulczynski, 1899 |
| Range: Portugal (Madeira) (van Helsdingen 2009.1). Global range: Madeira (Platnick 10.0). |
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| Mesiotelus mauritanicus Simon, 1909 |
| Range: France (Mainland), Italy (Sicily), Portugal (Mainland), Spain (Mainland) (van Helsdingen 2009.1). Global range: Mediterranean (Platnick 10.0). |
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| Mesiotelus tenellus (Thorell, 1875) |
| Range: Italy (Mainland) (van Helsdingen 2009.1). Global range: Italy (Platnick 10.0). |
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| Mesiotelus tenuissimus (L. Koch, 1866) |
| Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Crete), Greece (Cyclades), Greece (Mainland), Italy (Mainland), Italy (Sardinia), Italy (Sicily), Macedonia, Malta, Portugal (Mainland), Spain (Mainland), Ukraine (van Helsdingen 2009.1). Global range: Europe, North Africa, Turkmenistan (Platnick 10.0). |
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| Mesiotelus viridis (L. Koch, 1867) |
| Range: Greece (Cyclades), Greece (Dodecanese Islands), Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0). |
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| Genus: Sagana Thorell, 1875 |
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| Sagana rutilans Thorell, 1875 |
| Range: Austria, Bosnia/Herzegowina, Bulgaria, Croatia, Czech Republic, France (Mainland), Germany, Greece (Mainland), Hungary, Italy (Mainland), Macedonia, Netherlands, Poland, Romania, Slovakia, Slovenia, Switzerland, Yugoslavia (van Helsdingen 2009.1). Global range: Europe to Georgia (Platnick 10.0). |
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| Genus: Scotina Menge, 1873 |
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| Characters of genus: These are small spiders with six to ten pairs of ventral spines easily visible with a lens. The cephalothorax is darker than in the other genera of the family and shiny. The species are found at ground level among moss and detritus. The femora of the legs are light brown while more distal segments are darkened, particularly for the first pairs of legs. The two Danish species are easily separable by their markings. |
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| Scotina celans (Blackwall, 1841) |
| Description: The male has much smaller abdomen than the female, and is therefore overall smaller. The cephalothorax has characteristic light median and lateral bands, and may at distance resemble immature Lycosids of the genus Pardosa. The abdomen is reddish-brown, and the markings are typical of the Liocranidae, with grey chevrons. These are however, rather obscure for the male. Size: Female 4-5 mm; male 2.5-3 mm. Maturity: Males in autumn, females all year? Habitat: Damp moss, leaf litter and tussocks on juniper heathland, forest edges and forest clearings. Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Hungary, Ireland, Italy (Mainland), Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Russia (Eastern European), Russia (Kaliningrad Region), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe, Algeria, Russia (Platnick 10.0). |
Male. |
Female. |
Female. |
Female. |
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| Scotina gracilipes (Blackwall, 1859) |
| Description: Carapace with same markings as S. celans with median and lateral bands. However, these bands are much darker and the carapace appear uniform dark.Carapace is somewhat shinier than that of S. celans. The abdomen is also darker with obscure chevrons. The colouration of the legs are also generally similar to that of S. celans but the dark segments are darker, almost black in legs I and II, as well as shinier. Rear legs have annulated appearance. Size: 2.5-3.5 mm. Maturity: Males late summer and autumn, females throughout year. Habitat: Dry sandy places such as coastal and inland dunes and heathland. Range: Belgium, Croatia, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Ireland, Netherlands, Norway (Mainland), Poland, Russia (Kaliningrad Region), Slovenia, Sweden, Switzerland (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0). |
Female. |
Male. |
Female. |
Female. |
Female. |
Female. |
Female. |
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| Scotina occulta Kritscher, 1996 |
| Range: Malta (van Helsdingen 2009.1). Global range: Malta (Platnick 10.0). |
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| Scotina palliardii (L. Koch, 1881) |
| Range: Austria, Belgium, Czech Republic, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Latvia, Lithuania, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0). |
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