- filtered for Bosnia/Herzegowina
Images and Species Descriptions
Text and photographs © 2011 Jørgen Lissner
The Spiders of Europe and Greenland
Family: Linyphiidae (Line Weaving Spiders)
Biology: The Linyphiidae is the worlds second largest spider family encompassing ca. 4320 species in more than 570 genera. The highest diversity is found in the northern temperate regions. In these regions as well as in the arctic regions spiders of this family dominates the spider fauna. The jumping spiders (Salticidae) is an even larger family but they generally require warmer conditions than the linyphiids and for this reason they are less well represented in the cooler regions. Linyphiids are found worldwide in all terrestrial biotopes and is perhaps the most widely distributed spider family. They range in size from very small to medium (1-8.5 mm body size). Most species are found at ground level but they occupy a very wide array of habitats. The members of the family build a sheet web sometimes dome shaped. The web has no retreat and the spider always hang inverted below the sheet. Larger species in particular sometimes add irregular vertical snares acting both as sheet suspension strands and barrage balloon wires impeding the flight of insects. When insects strike the vertical snares they fall down on the sheet and the spider rush to bite the prey through the sheet web. Many species disperse by air and the phenomenon of ballooning is very noticeable in this family when very dense populations try to balloon at the same time. The spiders climb up high in the vegetation and point the spinners toward the sky. In this position they let out some strands of silk and eventually the wind will lift the spiders up in the air. Often they only manage to fly a short distance but they will keep trying sometimes resulting in a layer of shimmering silk covering the vegetation. Ballooning takes place usually in late summer. Ballooning may result in spiders literally raining from the sky. Erigone atra is a very common aeronaut in late summer in some areas and on several occasions I have experienced specimens landing in my hair at short intervals, for example while I was sitting in my garden.
Characters of family: The linyphiids belong to the group of ecribellate spider families having 8 eyes and 3 tarsal claws. The eyes are arranged in 2 rows of 4, usually heterogeneous in size with the anterior medials smaller than the rest. Frequently, the eyes are ringed with black, this being most noticeable in species with lighter coloured carapaces such as many species of the Linyphiinae subfamily. Some species adapted to dark habitats have the eyes much reduced, sometimes being very minute in size or only evidenced by pale markings under the integument (e.g. Porrhomma rosenhauri). The carapace is highly variable especially in the smaller species belonging to the Erigoninae subfamily. Males of this large subfamily frequently have the frontal region modified into strangely formed lobes or bear other types of protuberances some of which may have tufts of hairs. Some species have the carapace punctured with pits (see images of Lophomma punctatum). The males may also have sulci (grooves) running backwards from the posterior eyes. The chelicerae do not possess a lateral condyle (boss at base of chelicer). The outer side of the chelicerae have horizontal stridulating ridges visible in many species. Such ridges only occur scattered in other spider families (see for example images of ridges in Metellina stridulans of the Tetragnathidae). The labium is strongly rebordered as in the Nesticidae, Araneidae, and Tetragnathidae. The endites are usually parallel. Legs are slender and provided with spines. The number of spines on the legs is an important character for species identification when this is undertaken using the stereomicroscope. The abdomen is nearly always longer than wide sometimes with a pattern (Linyphiinae in particular) and sometimes mainly uniformly coloured, very often blackish (Erigoninae in particular). Some species posses an abdominal scutum as for example some members of the Ceratinella genus. The epigynes are variable, sometimes simple as in the Erigoninae or provided with a scapus as often seen in the Linyphiinae. The male palp often possesses an U-shaped paracymbium. The family was earlier divided in to two subfamilies, which sometimes were elevated to family status: the Linyphiidae and Erigonidae (also known as Micryphantidae). The Linyphiidae was characterized by not having tibial apophyses on the male palp, by having a claw on the female palp in most species, and by having two dorsal spines on tibia IV or if only one spine present there was one short spine on metatarsi I and II. The Erigonidae was characterized by having at least one tibial apophyses on the male palp, by lacking a claw on the female palp, and by having just a single dorsal spine on tibia IV and with the metatarsi spineless, or all spines lacking altogether. However, the family is now divided in to seven subfamilies, the Dubiaraneinae, Erigoninae, Ipainae, Linyphiinae, Micronetinae, Mynogleninae, and Stemonyphantinae. Consult recent literature or Wikipedia for lists of subfamily genera. See also Linyphiid Spiders Of The World by Andrei Tanasevitch and LinyGen: Linyphioid Genera of the World (Pimoidae and Linyphiidae) by Gustavo Hormiga, Dimitar Dimitrov, Jeremy A. Miller and Fernando Alvarez-Padilla.
Genus: Centromerus Dahl, 1886
Characters of genus: Legs fairly short and stout (Locket & Millidge 1953). Metatarsi I and II with a small dorsal spine. Metatarsus IV without a trichobothrium. Tibia I sometimes with a prolateral spine. Tibia IV with one or two dorsal spines. Epigyne with a scape in most species.
Centromerus subcaecus Kulczynski, 1914
Range: Austria, Bosnia/Herzegowina, Croatia, Germany, Italy (Mainland), Yugoslavia (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0).
Genus: Fageiella Kratochvl, 1934
Fageiella patellata (Kulczynski, 1913)
Range: Bosnia/Herzegowina, Yugoslavia (van Helsdingen 2009.1). Global range: Southeastern Europe (Platnick 10.0).
Genus: Lepthyphantes Menge, 1866
Biology: Due to the high number of species in this genus (formerly almost 400) many former Lepthyphantes species have been grouped in to new genera mainly based on small details in the copulatory organs. Saaristo & Tanasevitch (1996) limit the genus to only five species of which only two are European, but this is apparently not followed by Platnick.
Characters of genus: Medium to large linyphiids ranging form 2.5 to 4.5 mm body length (Saaristo & Tanasevitch 1996). Legs clearly annulated in European species. Tm IV without trichobothrium. Abdomen with a distinct pattern, usually composed of broad black transverse markings formed as bars, bands or chevrons on a grey background. Abdomens may be dotted with some whitish spots. Males are characterized by the sickle-shaped embolus with tight sulcus and large carina (not visible with a hand lens). Epigynes wit large scape arising from the inside of the epigynal cavity. Note that there is some disagreements on the delineation of the genus. The genus description here is based on a narrow conception as given by Saaristo & Tanasevitch (1996).
Lepthyphantes centromeroides Kulczynski, 1914
Range: Bosnia/Herzegowina, Bulgaria, Macedonia, Romania, Yugoslavia (van Helsdingen 2009.1). Global range: Balkans, Bulgaria, Romania (Platnick 10.0).
Genus: Palliduphantes Saaristo & Tanasevitch, 2001
Biology: Pale coloured spiders, mostly more or less cavernicolous in lifestyle. The genus incorporates a group of former Lepthyphantes species now reclassified to the new Palliduphantes genus based on synapomorphies of the secondary genital organs (Saaristo & Tanasevitch 2001). The males of the species share some special characters including similar shaped paracymbia which are large and tub-like. Also the lamella of the palps and the epigynes share similarities, the epigynes having a very characteristic appearance in dorsal view (see Saaristo & Tanasevitch 2001 for more details on diagnostic characters). The genus is subdivided in to eight species groups bases on the morphology of the secondary genital organs (Saaristo & Tanasevitch 2001)
Characters of genus: Small to medium sized linyphiids having body lengths ranging from 1.30-2.95 mm, but species larger than 2.5 mm are few (Saaristo & Tanasevitch 2001). Pale coloured spiders, the cephalothorax and appendages range in colour from pale yellow to pale orange and the unicoloured abdomen from pale yellow to grey or greyish brown, sometimes with a greenish tinge or a faint pattern of transverse stripes or chevrons. Legs with few spines. Metatarsus IV is without a trichobotrium.
Palliduphantes istrianus (Kulczynski, 1914)
Range: Bosnia/Herzegowina, Bulgaria, Croatia, Greece (Mainland), Hungary, Italy (Mainland), Romania, Slovenia, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Eastern Europe (Platnick 10.0).
Palliduphantes pallidus (O. P.-Cambridge, 1871)
Range: Austria, Belarus, Belgium, Bosnia/Herzegowina, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Hungary, Iceland, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
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Male.
Male.
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Female.
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Female.
Palliduphantes spelaeorum (Kulczynski, 1914)
Range: Bosnia/Herzegowina, Bulgaria, Croatia, Greece (Mainland), Macedonia, Yugoslavia (van Helsdingen 2009.1). Global range: Balkans, Bulgaria, Greece (Platnick 10.0).
Genus: Porrhomma Simon, 1884
Biology: A fairly homogenous group of small spiders which are difficult to identify due to similarity of species and rather great intraspecific variation. Some species are adapted morphologically to troglodytic or subterranean lifestyle, e.g. with eyes distinctly reduced or with paler pigmentation. Depending on habitat, troglodytic species exhibit a great degree of plasticity in morphological adaptations (even leg spination varies) which further complicates identification. The species of Porrhomma are mainly found in the temperate zone of the northern Hemisphere, with most species in the Palaearctic region.
Characters of genus: There is no recent revision of the genus available. The description here is based on Locket & Millidge (1953), and Borges & Wunderlich (2008). Ocular area with some forward projecting hairs, most pronounced in males. Eyes are variable in size, in some species minute or absent. Metatarsus IV without a trichobothrium. The species differ in leg spination, a character that is useful for grouping the species aiding identification. Male palp with no tibial apophysis. Some species possess stridulating files on coxa I while these are reduced or absent in others. The latter case is considered a derived (apomorphic) character of the genus. Tm I of British species range between 0.3-0.64 (Roberts 1987).
Porrhomma convexum (Westring, 1851)
Description: A fairly large Porrhomma species. Carapace yellow-brown to blackish brown. Ocular area with forward projecting hairs. Abdomen greyish black. Legs yellow-brown. Femur I with one or two dorsal spines and without prolateral spine. All metatarsi spineless. Tm I 0.4-0.49 (Roberts 1987). Size: 2.2-3.0 mm. Range: Austria, Belarus, Belgium, Bosnia/Herzegowina, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Faroe Islands, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Hungary, Iceland, Ireland, Italy (Mainland), Italy (Sardinia), Liechtenstein, Macedonia, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Also known from Greenland (Masurik in prep.). Global range: Palearctic (Platnick 10.0).
Male.
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Genus: Syedra Simon, 1884
Syedra caporiaccoi Kolosvry, 1938
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Genus: Tenuiphantes Saaristo & Tanasevitch, 1996
Biology: Previously, members of this genus were placed in the species rich genus of Lepthyphantes. A homogenous group of species have now been transferred to the Tenuiphantes genus which is characterised by their copulatory organs. Markings can be quite variable within species, nevertheless they are sometimes useful for separating the species.
Characters of genus: Small to large linyphiids ranging from 1.7-4.1 mm body length (Saaristo & Tanasevitch 1996). Males are characterized by having a sinuous embolus, often with a dentigerous protrusion at about halfway while females are characterized by having the proscapus bordered at either side by a lateral wing-like extension of the median part of the scapus (Saaristo & Tanasevitch 1996). Epigynes of T. alacris females also characterized by strongly developed lateral teeth. Paracymbium with 0-3 teeth. The species differ in leg spination. Metatarsus without a trichobothrium except in T. retezaticus (endemic to Romania). Most species have uniformly coloured legs and a dark dorsal pattern on the abdomen usually composed of broad black transverse bars on a brownish background. Bars sometimes reduced to paired dots combined by thin black lines. Dorsal pattern is usually more obscure in males. Legs fairly long and with long spines.
Tenuiphantes tenebricola (Wider, 1834)
Range: Austria, Belarus, Belgium, Bosnia/Herzegowina, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Genus: Troglohyphantes Joseph, 1881
Troglohyphantes affinis (Kulczynski, 1914)
Range: Bosnia/Herzegowina, Croatia, Spain (Mainland) (van Helsdingen 2009.1). Global range: Croatia, Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes brevipes Deeleman-Reinhold, 1978
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes dekkingae Deeleman-Reinhold, 1978
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes dekkingae pauciaculeatus Deeleman-Reinhold, 1978
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes fallax Deeleman-Reinhold, 1978
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes fugax (Kulczynski, 1914)
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina8 (Platnick 10.0).
Troglohyphantes hadzii Kratochvíl, 1934
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes milleri (Kratochvíl, 1948)
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes montanus Absolon & Kratochvíl, 1932
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes pugnax Deeleman-Reinhold, 1978
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes salax (Kulczynski, 1914)
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes svilajensis (Kratochvíl, 1948)
Range: Bosnia/Herzegowina, Croatia (van Helsdingen 2009.1). Global range: Croatia (Platnick 10.0).
Troglohyphantes svilajensis bosnicus (Kratochvíl, 1948)
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes troglodytes (Kulczynski, 1914)
Range: Bosnia/Herzegowina, Yugoslavia (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).
Troglohyphantes wiebesi Deeleman-Reinhold, 1978
Range: Bosnia/Herzegowina (van Helsdingen 2009.1). Global range: Bosnia-Hercegovina (Platnick 10.0).