- filtered for Greece (Mainland)
Images and Species Descriptions
Text and photographs © 2011 Jørgen Lissner
The Spiders of Europe and Greenland
Family: Linyphiidae (Line Weaving Spiders)
Biology: The Linyphiidae is the worlds second largest spider family encompassing ca. 4320 species in more than 570 genera. The highest diversity is found in the northern temperate regions. In these regions as well as in the arctic regions spiders of this family dominates the spider fauna. The jumping spiders (Salticidae) is an even larger family but they generally require warmer conditions than the linyphiids and for this reason they are less well represented in the cooler regions. Linyphiids are found worldwide in all terrestrial biotopes and is perhaps the most widely distributed spider family. They range in size from very small to medium (1-8.5 mm body size). Most species are found at ground level but they occupy a very wide array of habitats. The members of the family build a sheet web sometimes dome shaped. The web has no retreat and the spider always hang inverted below the sheet. Larger species in particular sometimes add irregular vertical snares acting both as sheet suspension strands and barrage balloon wires impeding the flight of insects. When insects strike the vertical snares they fall down on the sheet and the spider rush to bite the prey through the sheet web. Many species disperse by air and the phenomenon of ballooning is very noticeable in this family when very dense populations try to balloon at the same time. The spiders climb up high in the vegetation and point the spinners toward the sky. In this position they let out some strands of silk and eventually the wind will lift the spiders up in the air. Often they only manage to fly a short distance but they will keep trying sometimes resulting in a layer of shimmering silk covering the vegetation. Ballooning takes place usually in late summer. Ballooning may result in spiders literally raining from the sky. Erigone atra is a very common aeronaut in late summer in some areas and on several occasions I have experienced specimens landing in my hair at short intervals, for example while I was sitting in my garden.
Characters of family: The linyphiids belong to the group of ecribellate spider families having 8 eyes and 3 tarsal claws. The eyes are arranged in 2 rows of 4, usually heterogeneous in size with the anterior medials smaller than the rest. Frequently, the eyes are ringed with black, this being most noticeable in species with lighter coloured carapaces such as many species of the Linyphiinae subfamily. Some species adapted to dark habitats have the eyes much reduced, sometimes being very minute in size or only evidenced by pale markings under the integument (e.g. Porrhomma rosenhauri). The carapace is highly variable especially in the smaller species belonging to the Erigoninae subfamily. Males of this large subfamily frequently have the frontal region modified into strangely formed lobes or bear other types of protuberances some of which may have tufts of hairs. Some species have the carapace punctured with pits (see images of Lophomma punctatum). The males may also have sulci (grooves) running backwards from the posterior eyes. The chelicerae do not possess a lateral condyle (boss at base of chelicer). The outer side of the chelicerae have horizontal stridulating ridges visible in many species. Such ridges only occur scattered in other spider families (see for example images of ridges in Metellina stridulans of the Tetragnathidae). The labium is strongly rebordered as in the Nesticidae, Araneidae, and Tetragnathidae. The endites are usually parallel. Legs are slender and provided with spines. The number of spines on the legs is an important character for species identification when this is undertaken using the stereomicroscope. The abdomen is nearly always longer than wide sometimes with a pattern (Linyphiinae in particular) and sometimes mainly uniformly coloured, very often blackish (Erigoninae in particular). Some species posses an abdominal scutum as for example some members of the Ceratinella genus. The epigynes are variable, sometimes simple as in the Erigoninae or provided with a scapus as often seen in the Linyphiinae. The male palp often possesses an U-shaped paracymbium. The family was earlier divided in to two subfamilies, which sometimes were elevated to family status: the Linyphiidae and Erigonidae (also known as Micryphantidae). The Linyphiidae was characterized by not having tibial apophyses on the male palp, by having a claw on the female palp in most species, and by having two dorsal spines on tibia IV or if only one spine present there was one short spine on metatarsi I and II. The Erigonidae was characterized by having at least one tibial apophyses on the male palp, by lacking a claw on the female palp, and by having just a single dorsal spine on tibia IV and with the metatarsi spineless, or all spines lacking altogether. However, the family is now divided in to seven subfamilies, the Dubiaraneinae, Erigoninae, Ipainae, Linyphiinae, Micronetinae, Mynogleninae, and Stemonyphantinae. Consult recent literature or Wikipedia for lists of subfamily genera. See also Linyphiid Spiders Of The World by Andrei Tanasevitch and LinyGen: Linyphioid Genera of the World (Pimoidae and Linyphiidae) by Gustavo Hormiga, Dimitar Dimitrov, Jeremy A. Miller and Fernando Alvarez-Padilla.
Genus: Acartauchenius Simon, 1884
Acartauchenius scurrilis (O. P.-Cambridge, 1872)
Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Lithuania, Macedonia, Netherlands, Poland, Romania, Russia (Eastern European), Russia (Southern European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Eggsack visible behind curtain of silk. It contained only four eggs which were relatively large compared to the spider.
Eggsack placed in cavity on underside of stone.
Female.
Female.
Female.
Female.
Female.
Male.
Male.
Male.
Male.
Male.
Habitat.
Genus: Agyneta Hull, 1911
Biology: Agyneta share some morphological traits with Meioneta including tracheae extending in to the cephalothorax as well as similar structure of the secondary genital organs. This has resulted in synonymization and transfer of species from Meioneta to Agyneta (see for example Saaristo 1973, Roberts 1987 and Saaristo & Koponen 1998). However, the present view is to consider this incorrect. Millidge (1977) point out that the two genera most likely constitute two phylogenetic lines due to non-overlapping Tm I range, a view that has not yet been proven wrong (The World Spider Catalog by Platnick). There are a few other differences between the genera. Meioneta lacks a trichobothrium on metatarsus IV and possess different cheliceral teeth than Agyneta. Some Meioneta species possess lateral spines on tibia I and II. Members of Agyneta have only dorsal spines on tibia I-II. From a practical point of view, it is convenient to uphold the two genera due to the fairly high number of species, totalling 30 in Europe alone. Species rich genera are unhandy in terms of species identification. Identification of single females in the two genera may be difficult as the members have very similar epigynes, and it is sometimes stated that a female can be identified only from its habitat and association with the male (Saaristo & Koponen 1998). However, some help may be obtained from the degree of swelling and blackening of the female palp which differs among species.
Characters of genus: Posterior eyes of approximately similar size (Locket & Millidge, 1953). Tm I = 0.65-0.9. With a trichobothrium on metatarsus IV. Lateral spines on Tibia I and II absent.
Agyneta subtilis (O. P.-Cambridge, 1863)
Description: Carapace brown. Abdomen grey to black. Legs orange brown or brown with tibiae I and metatarsus I darker. Female palp strongly swollen. Male palp elevated dorsally, appearing subquadratic in lateral view. Size: Female 2.0-2.7 mm; male 2.0-2.5 mm. Range: Austria, Belarus, Belgium, Czech Republic, Denmark, Estonia, Faroe Islands, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Ireland, Italy (Mainland), Latvia, Lithuania, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Also known from Greenland (Masurik in prep.). Global range: Palearctic (Platnick 10.0).
Genus: Araeoncus Simon, 1884
Araeoncus humilis (Blackwall, 1841)
male 1.5 mm. Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Malta, Netherlands, Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic, New Zealand (Platnick 10.0).
Male.
Female.
Female.
Female.
Female.
Male.
Male.
Female.
Female.
Genus: Bathyphantes Menge, 1866
Characters of genus: Palp of female with tarsal claw. Legs rather long and slender. All tibiae with two dorsal spines, metatarsi spineless (Locket & Millidge 1953).
Bathyphantes parvulus (Westring, 1851)
Range: Austria, Belgium, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Ireland, Italy (Mainland), Lithuania, Netherlands, Norway (Mainland), Poland, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (NW. European), Slovakia, Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Genus: Bolyphantes C. L. Koch, 1837
Characters of genus: Fairly large linyphiidsof the subfamily Micronetinae having body lengths up to 5 mm (Saaristo & Tanasesevitch 2000). In some species the male carapace is elevated in the ocular area which is furnished with numerous forward directed bristles. Carapace only somewhat raised in female. Abdomen with pattern. Metatarsus IV without a trichobothrium. Legs long and slender and well armed with spines. Femora without dorsal spines. Tibiae and metatarsi with numerous spines. Tm I 0.15-0.18 (Saaristo & Tanasesevitch 2000). Abdomen with pattern. Males have a stout spine on the palpal patella.
Bolyphantes lamellaris Tanasevitch, 1990
Range: Greece (Mainland), Italy (Mainland) (van Helsdingen 2009.1). Global range: Italy, Greece, Russia (Platnick 10.0).
Genus: Centromerus Dahl, 1886
Characters of genus: Legs fairly short and stout (Locket & Millidge 1953). Metatarsi I and II with a small dorsal spine. Metatarsus IV without a trichobothrium. Tibia I sometimes with a prolateral spine. Tibia IV with one or two dorsal spines. Epigyne with a scape in most species.
Centromerus sylvaticus (Blackwall, 1841)
Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Balearic Islands), Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Female.
Female.
Male.
Male.
Male.
Male.
Male.
Genus: Ceratinella Emerton, 1882
Characters of genus: Compact dark spiders with coriaceous abdomens (Locket & Millidge 1953). The male carapace is slightly domed in head region. The legs are short and robust with metatarsi equal or slightly longer than tarsi. Tibial spines very short and resemble hairs. The abdomen is globular in females, less so in males. A dorsal scutum is present in many species.
Ceratinella brevis (Wider, 1834)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia?, Liechtenstein, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Female.
Male.
Male.
Male.
Male.
Male.
Ceratinella scabrosa (O. P.-Cambridge, 1871)
Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Genus: Cresmatoneta Simon, 1929
Cresmatoneta mutinensis (Canestrini, 1868)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Greece (Mainland), Italy (Mainland), Macedonia, Romania, Slovenia, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Genus: Dicymbium Menge, 1868
Dicymbium nigrum (Blackwall, 1834)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Dicymbium tibiale (Blackwall, 1836)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Ireland, Latvia, Liechtenstein, Lithuania, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Male. Note the swollen Tibia I.
Male.
Male with mite.
Male with mite.
Female.
Female.
Female.
Female.
Female.
Male.
Male.
Male.
Genus: Diplocephalus Bertkau, 1883
Characters of genus: Male head elevated in to a variety of lobes (Locket & Millidge 1953). Tibiae I-II with two spines, III-IV with just one spine (spines sometimes reduced or absent in males). Metatarsi longer than tarsi: leg I: 1.2-1.3 times, leg IV: 1.5-1.6 times (Locket & Millidge 1953).
Diplocephalus alpinus (O. P.-Cambridge, 1872)
Range: Austria, Croatia, Greece (Mainland), Hungary, Italy (Mainland), Liechtenstein, Romania, Russia (Central European), Slovenia, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Italy, Central Europe to Russia (Platnick 10.0).
Diplocephalus cristatus (Blackwall, 1833)
Description: Carapace brown to greyish black. Striae not visible in live specimens. Male head elevated anteriorly divided by a transverse groove. Eyes fairly large in female and posterior medials are about 0.7 diameter apart and about 0.5 diameter apart from laterals. Eyes are smaller in males and due to the elevation of the head farther apart (2-3 diameter between posterior medials and laterals). Abdomen is grey to black. Legs are grey (recently moulted specimens?) to reddish brown. Size: Female 1.8-2.5 mm; male 1.7-2.1 mm. Range: Austria, Belarus, Belgium, Croatia, Czech Republic, Denmark, Estonia, Faroe Islands, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Iceland, Ireland, Italy (Mainland), Liechtenstein, Lithuania, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic, New Zealand, Falkland Is (Platnick 10.0).
Male.
Male.
Male.
Female.
Female.
Female.
Female.
Female.
Female.
Female.
Male.
Male.
Male.
Male.
Male.
Diplocephalus graecus (O. P.-Cambridge, 1872)
Range: Belgium, Bulgaria, France (Corsica), France (Mainland), Great Britain (Mainland), Greece (Mainland), Italy (Mainland), Portugal (Mainland), Romania, Spain (Balearic Islands), Spain (Mainland) (van Helsdingen 2009.1). Global range: Southern, Central Europe, North Africa (Platnick 10.0).
Diplocephalus permixtus (O. P.-Cambridge, 1871)
Description: Carapace brown to greyish black. Striae not visible in live specimens. Male head elevated anteriorly divided by a transverse groove. Male carapace rather elongate and projecting considerably over chelicerae. In the female posterior medials are about 0.8 diameter apart and about 0.5 diameter apart from laterals. Eyes are smaller in males and due to the elevation of the head farther apart (2.5 diameter between posterior medials and laterals). Abdomen is grey to black. Legs are brown to yellow brown. Tibia I spineless in males. Size: Female 1.6-1.9 mm; male 1.5-1.9 mm. Range: Austria, Belgium, Bulgaria, Czech Republic, Denmark, Faroe Islands, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Iceland, Ireland, Lithuania, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (NW. European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Female.
Female.
Male.
Male.
Male.
Female.
Female.
Diplocephalus picinus (Blackwall, 1841)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Female.
Diplocephalus turcicus Brignoli, 1972
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece, Turkey (Platnick 10.0).
Genus: Diplostyla Emerton, 1882
Diplostyla concolor (Wider, 1834)
Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Madeira), Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Female.
Female.
Male.
Male.
Genus: Entelecara Simon, 1884
Characters of genus: Most species are rather similar in general appearance. Males have the head region domed. Female tibiae I-II with two spines, III-IV with just one. In males spines are much reduced, usually absent altogether from tibiae I-II (Locket & Millidge 1953). Metatarsus IV with a trichobothrium in most species. Male palp with two prominent tibial apophyses.
Entelecara acuminata (Wider, 1834)
Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Latvia, Lithuania, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Male.
Male.
Male.
Genus: Erigone Audouin, 1826
Characters of genus: Edges of carapace strongly dentate in males, less so in females. Male head domed, but not raised in to a lobe. The chelicers are robust and furnished with warts and teeth anteriorly being more strongly developed in males than in females. Tibiae I-III with two spines, IV with just one. Metatarsi slightly longer than tarsi: leg I: 1.3-1.4 times, leg IV: ca.1.6 times (Locket & Millidge 1953). Male palp with characteristic large ventral patellar apophysis at the distal end. Palpal femur often with ventral knobs and teeth along length.
Erigone dentipalpis (Wider, 1834)
Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Greece (North Aegean Islands), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Azores), Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Balearic Islands), Spain (Canary Islands), Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Genus: Erigonoplus Simon, 1884
Erigonoplus setosus Wunderlich, 1995
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Croatia, Greece (Platnick 10.0).
Genus: Frontinellina van Helsdingen, 1969
Frontinellina frutetorum (C. L. Koch, 1834)
Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Cyprus, Czech Republic, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Crete), Greece (Mainland), Greece (North Aegean Islands), Hungary, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Liechtenstein, Macedonia, Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Subadult female.
Subadult female.
Subadult female.
Subadult female.
Subadult female.
Subadult female.
Male.
Male.
Male.
Female.
Female.
Female.
Male.
Genus: Gnathonarium Karsch, 1881
Gnathonarium dentatum (Wider, 1834)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Cyclades), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Italy (Sicily), Latvia, Liechtenstein, Lithuania, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male, characteristic palps shaped like a droplet.
Male.
Male.
Male.
Male.
Male palp.
Male.
Genus: Gonatium Menge, 1868
Biology: This genus has been reviewed by Millidge (1981). It is possible to recognise the members by the following combination of somatic characters: leg spination, presence of a tricobothrium on metatarsus IV, value of Tm I, and by the way the tarsal claws are pectinated (see description below). The genus is, however, also defined by the structure of the genitals which share characters thought to be derived (see Millidge (1981) for details).
Characters of genus: The species range in size from 2.0-3.7 mm. The cephalothorax and legs are bright orange, orange red or reddish brown. The male head carries no lobe but is slightly raised. Sternum at least as wide as long with coxae IV widely separated. Legs fairly long and slender. Leg spines are short, shorter and weaker in males than in females. Males have tibia I and to a lesser degree tibia II curved and swollen distally. They are furnished ventrally with many long hairs or bristles. Also metatarsi I and II and femora I and II are provided with many short spines or bristles ventrally. There is a single spine on each tibia in both sexes. All metatarsi with a trichobothrium. Tm I range from 0.75-0.95 (Millidge 1981). The tarsal claws are pectinated, consisting of narrow, needle-like teeth. The pectination is different in almost all other groups of spiders with pectinate claws (Millidge 1981). Abdomen globular, often reddish with four sigilla dorsally. There is no scutum. The male palpal femur is swollen in some species. The epigynes of all Gonatium species have the same general appearance, and depending on the number of species at a given locality specimens may be identifiable with a lens.
Gonatium hilare (Thorell, 1875)
Range: Andorra, Austria, Belgium, Bulgaria, Croatia, Czech Republic, France (Corsica), France (Mainland), Germany, Greece (Mainland), Hungary, Italy (Mainland), Italy (Sicily), Liechtenstein, Macedonia, Romania, Slovakia, Slovenia, Spain (Mainland), Switzerland (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Gonatium rubellum (Blackwall, 1841)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Female.
Female.
Female.
Genus: Gongylidiellum Simon, 1884
Characters of genus: The genus contains mostly small species less than 2 mm body length. Genus characters are not visible with a hand lens but require a stereomicroscope. Male head not elevated. Apparently, males use sound production in courtship as the branchial opercula are provided with stridulating ridges opposing a tooth-like outgrowth on coxae IV. Metatarsus IV without a trichobothrium. Tibiae I-II with two spines, III-IV with just one.
Gongylidiellum murcidum Simon, 1884
Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Lithuania, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Slovakia, Slovenia?, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Genus: Gongylidium Menge, 1868
Gongylidium rufipes (Linnaeus, 1758)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Male.
Male.
Male.
Male.
Male.
Female.
Female.
Female.
Female.
Female.
Genus: Incestophantes Tanasevitch, 1992
Incestophantes frigidus (Simon, 1884)
Range: France (Mainland), Greece (Mainland)?, Italy (Mainland), Liechtenstein, Macedonia, Slovenia?, Switzerland (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0).
Genus: Lepthyphantes Menge, 1866
Biology: Due to the high number of species in this genus (formerly almost 400) many former Lepthyphantes species have been grouped in to new genera mainly based on small details in the copulatory organs. Saaristo & Tanasevitch (1996) limit the genus to only five species of which only two are European, but this is apparently not followed by Platnick.
Characters of genus: Medium to large linyphiids ranging form 2.5 to 4.5 mm body length (Saaristo & Tanasevitch 1996). Legs clearly annulated in European species. Tm IV without trichobothrium. Abdomen with a distinct pattern, usually composed of broad black transverse markings formed as bars, bands or chevrons on a grey background. Abdomens may be dotted with some whitish spots. Males are characterized by the sickle-shaped embolus with tight sulcus and large carina (not visible with a hand lens). Epigynes wit large scape arising from the inside of the epigynal cavity. Note that there is some disagreements on the delineation of the genus. The genus description here is based on a narrow conception as given by Saaristo & Tanasevitch (1996).
Lepthyphantes corfuensis Wunderlich, 1995
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Lepthyphantes leprosus (Ohlert, 1865) - House line-weaver
Description: Carapace yellow-brown to brown with darker striae (Locket & Millidge 1953). Legs similarly coloured but with blackish annulations and relatively long spines. Abdomen brownish with transverse chevron-like black bars and with black sides. Brown areas with white, mostly small dots of varying forms and sizes. Tm I = ca. 0.13. Etymology: the latin species name "leprosus" means appearing decayed, having the appearance of infection by leprosy (Wiktionary). Size: Female 2.4-3.8 mm; male 2.4-3.5 mm. Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Faroe Islands?, Finland, France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Iceland, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic, Chile (Platnick 10.0).
Female.
Female.
Female.
Female.
Female.
Male.
Male.
Male.
Male.
Lepthyphantes magnesiae Brignoli, 1979
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Lepthyphantes minutus (Blackwall, 1833)
Description: Lepthyphantes minutus is one of the larger Lepthyphantes species despite that the species name indicate the opposite. The explanation is that the species earlier was assigned to the Linyphia genus which consists of relatively larger species. The abdomen has a beatifully golden colouration with ligther spots. Habitat: The species is common in woods where i build a sheet web at the bases of tree trunks, under logs and in holes of trees. Range: Austria, Belgium, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Male.
Male.
Male.
Male.
Female, venter.
Female.
Female.
Female.
Genus: Linyphia Latreille, 1804
Linyphia mimonti Simon, 1884
Range: Greece (Mainland), Italy (Mainland) (van Helsdingen 2009.1). Global range: Italy, Greece, Lebanon (Platnick 10.0).
Linyphia triangularis (Clerck, 1757)
Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Balearic Islands), Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic, introduced in USA (Platnick 10.0).
Female.
Male.
Male.
Male.
Female.
Female.
Male.
Male.
Male.
Male.
Female.
Female venter.
Female.
Female.
Female.
Genus: Maculoncus Wunderlich, 1995
Maculoncus parvipalpus Wunderlich, 1995
Range: Greece (Cyclades), Greece (Mainland), Greece (North Aegean Islands) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Genus: Maso Simon, 1884
Maso gallicus Simon, 1894
Range: Belgium, Bulgaria, Czech Republic, Estonia, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Italy (Mainland), Italy (Sardinia), Liechtenstein, Netherlands, Poland, Romania, Russia (Central European), Spain (Mainland), Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe to Azerbaijan (Platnick 10.0).
Female.
Female.
Genus: Mecopisthes Simon, 1926
Mecopisthes silus (O. P.-Cambridge, 1872)
Range: Austria, Croatia, Czech Republic, France (Corsica), France (Mainland), Germany, Greece (Mainland), Hungary, Italy (Mainland), Liechtenstein, Macedonia, Poland, Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Slovenia, Switzerland (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0).
Genus: Megalepthyphantes Wunderlich, 1994
Megalepthyphantes collinus (L. Koch, 1872)
Range: Austria, Bulgaria, Czech Republic, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Macedonia, Poland, Portugal (Mainland), Romania, Slovakia, Slovenia?, Spain (Mainland), Switzerland, Ukraine?, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Megalepthyphantes lydiae Wunderlich, 1994
Range: Greece (Cyclades), Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Male.
Male.
Male.
Female.
Female.
Female.
Megalepthyphantes nebulosus (Sundevall, 1830)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Lithuania, Macedonia, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Male.
Female.
Female.
Female.
Female.
Female.
Female abdominal markings.
Female.
Juvenile hanging in web in compost bin.
Female in sheet web.
Male.
Male abdominal markings.
Male.
Male palp.
Genus: Meioneta Hull, 1920
Characters of genus: Posterior eyes of approximately similar size or medians larger than laterals (Locket & Millidge 1953). Tm I = ca. 0.20-0.30. Some species possess lateral spines on Tibia I and II. Without a trichobothrium on metatarsus IV. Meioneta share some morphological traits with Agyneta (see genus description for Agyneta).
Meioneta fuscipalpa (C. L. Koch, 1836)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Estonia, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Lithuania, Macedonia, Moldova, Norway (Mainland), Poland, Portugal (Azores), Portugal (Madeira), Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (NW. European), Slovakia, Spain (Canary Islands), Spain (Mainland), Sweden?, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Palp and epigyne.
Meioneta punctata (Wunderlich, 1995)
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Meioneta simplicitarsis (Simon, 1884)
Range: Austria, Belgium, Bulgaria, Czech Republic, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Romania, Russia (Central European), Russia (Eastern European), Slovakia, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe, Russia, Kazakhstan (Platnick 10.0).
Genus: Metopobactrus Simon, 1884
Metopobactrus prominulus (O. P.-Cambridge, 1872)
Range: Austria, Belarus, Belgium, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Liechtenstein, Lithuania, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Also known from Greenland (Masurik in prep.). Global range: Holarctic (Platnick 10.0).
Female.
Female.
Male.
Male.
Male.
Female.
Female.
Male.
Male.
Genus: Microlinyphia Gerhardt, 1928
Characters of genus: Medium sized spiders 2.8-4.6 mm, larger species are found outside Europe (van Helsdingen 1970). This species displays a large sexual dimorphism. Cephalothorax of male is longer and narrower than in female. Abdomen tubular in male and usually dark while more oblong and lighter coloured in females. Posterior medial eyes on black tubercles. Lateral eyes contiguous. Legs long and slender, most noticeable in males. Embolus long and thread-like in European species, easily visible with a lens. The chelicerae in males are long, more than half the length of the cephalothorax and inclined somewhat backwards. The epigyne is small, inconspicuous and consists of an arch anterior to the openings and a small scape-like protrusion. The arch is dark and barely more sclerotized than surrounding area.
Microlinyphia pusilla (Sundevall, 1830)
Description: Male: Carapace brown to dark brown. Legs yellow-brown without annulations but with blackish streaks on basal half of femora in dark specimens. Abdomen tubular, dark brown with one pair of white spots anteriorly (rarely two pairs). The male palp has a large looped embolus which in lateral view encircle an area about twice as big as that covered by the palpal organs. Female: Cephalothorax less elongate than in male and brown. Abdomen is shorter and higher compared to males, ovoid in dorsal view. Ventral surface of lateral and posterior surface as well as ventral surface dark brown or blackish. Dorsally light coloured with a broad dark brown median band broken posteriorly in dark spots, the first being diamond shaped. Size: Female 3-5 mm; male 3-4 mm. Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Faroe Islands, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Crete), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Malta, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Female.
Male.
Male.
Male.
Female.
Female.
Female.
Female.
Female.
Female.
Male.
Male, note large hooped embolus of palp.
Male.
Female.
Female.
Female.
Genus: Neriene Blackwall, 1833
Neriene furtiva (O. P.-Cambridge, 1871)
Range: Andorra, Belgium, Bulgaria, Croatia, Czech Republic, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sardinia), Moldova, Netherlands, Poland, Portugal (Mainland), Romania, Russia (Eastern European), Russia (NW. European), Slovakia, Slovenia, Spain (Balearic Islands), Spain (Mainland), Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe, North Africa, Russia, Ukraine (Platnick 10.0).
Female.
Female.
Female.
Male.
Male.
Genus: Oedothorax Bertkau, in Förster & Bertkau, 1883
Oedothorax apicatus (Blackwall, 1850)
Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Moldova, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Male.
Female.
Female.
Male.
Female.
Female.
Female.
Female.
Female.
Male.
Male.
Male.
Male.
Male.
Male.
Genus: Palliduphantes Saaristo & Tanasevitch, 2001
Biology: Pale coloured spiders, mostly more or less cavernicolous in lifestyle. The genus incorporates a group of former Lepthyphantes species now reclassified to the new Palliduphantes genus based on synapomorphies of the secondary genital organs (Saaristo & Tanasevitch 2001). The males of the species share some special characters including similar shaped paracymbia which are large and tub-like. Also the lamella of the palps and the epigynes share similarities, the epigynes having a very characteristic appearance in dorsal view (see Saaristo & Tanasevitch 2001 for more details on diagnostic characters). The genus is subdivided in to eight species groups bases on the morphology of the secondary genital organs (Saaristo & Tanasevitch 2001)
Characters of genus: Small to medium sized linyphiids having body lengths ranging from 1.30-2.95 mm, but species larger than 2.5 mm are few (Saaristo & Tanasevitch 2001). Pale coloured spiders, the cephalothorax and appendages range in colour from pale yellow to pale orange and the unicoloured abdomen from pale yellow to grey or greyish brown, sometimes with a greenish tinge or a faint pattern of transverse stripes or chevrons. Legs with few spines. Metatarsus IV is without a trichobotrium.
Palliduphantes byzantinus (Fage, 1931)
Range: Bulgaria, Greece (Crete), Greece (Cyclades), Greece (Dodecanese Islands), Greece (Mainland), Macedonia, Romania (van Helsdingen 2009.1). Global range: Bulgaria, Greece, Turkey (Platnick 10.0).
Palliduphantes epaminondae (Brignoli, 1979)
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Palliduphantes istrianus (Kulczynski, 1914)
Range: Bosnia/Herzegowina, Bulgaria, Croatia, Greece (Mainland), Hungary, Italy (Mainland), Romania, Slovenia, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Eastern Europe (Platnick 10.0).
Palliduphantes spelaeorum (Kulczynski, 1914)
Range: Bosnia/Herzegowina, Bulgaria, Croatia, Greece (Mainland), Macedonia, Yugoslavia (van Helsdingen 2009.1). Global range: Balkans, Bulgaria, Greece (Platnick 10.0).
Genus: Pelecopsis Simon, 1864
Pelecopsis krausi Wunderlich, 1980
Range: Bulgaria, Greece (Mainland), Italy (Mainland), Macedonia (van Helsdingen 2009.1). Global range: Balkans to Armenia (Platnick 10.0).
Pelecopsis parallela (Wider, 1834)
Range: Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Portugal (Azores), Romania, Russia (Central European), Russia (Eastern European), Russia (Kaliningrad Region), Russia (Northern European), Russia (Novaya Zemlya), Russia (Southern European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Also known from Greenland (Masurik in prep.). Global range: Palearctic (Platnick 10.0).
Male.
Male.
Female.
Male.
Male.
Male.
Male.
Female.
Female.
Genus: Pocadicnemis Simon, 1884
Pocadicnemis juncea Locket & Millidge, 1953
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France (Mainland), Germany, Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Ireland, Italy (Mainland), Macedonia, Netherlands, Poland, Romania, Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Genus: Prinerigone Millidge, 1988
Prinerigone vagans (Audouin, 1826)
Range: Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sardinia), Italy (Sicily), Macedonia, Moldova, Netherlands, Portugal (Azores), Portugal (Madeira), Portugal (Mainland), Romania, Russia (Southern European), Slovakia, Spain (Balearic Islands), Spain (Canary Islands), Spain (Mainland), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Old World (Platnick 10.0).
Female.
Female.
Male.
Male.
Female.
Female.
Genus: Sauron Eskov, 1995
Sauron rayi (Simon, 1881)
Range: Belgium, Bulgaria, Croatia, France (Mainland), Germany, Greece (Mainland), Italy (Mainland), Poland, Romania, Slovakia, Slovenia (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0).
Genus: Silometopus Simon, 1926
Biology: Some species tolerates flooding with sea water (Nentwig W, Blick T, Gloor D, Hänggi A, Kropf C: Araneae - Spiders of Europe).
Characters of genus: The species in this genus are rather similar in general appearance and in their genitalia (Roberts 1985). Small species, body length range from 1.0- 2.2 mm. Tibia I-IV with one dorsal spines, usually weak or even absent in most males (Roberts 1985). Sometimes Tibia I-II are without dorsal spines (Nentwig W, Blick T, Gloor D, Hänggi A, Kropf C: Araneae - Spiders of Europe - Key to Linyphiidae). Metatarsus IV without a trichobothrium. Metatarsi not much longer than tarsi with Mt I/T1 ca. 1.1-1.2 (Locket & Millidge 1953). Head of males very slightly to slightly domed behind the eyes depending on species. Abdomen with four dark-brown dots (sigilla) which become reddish in alcohol preserved specimens.
Silometopus reussi (Thorell, 1871)
Range: Austria, Belgium, Bulgaria, Czech Republic, Denmark, Finland, France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Lithuania, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Spain (Mainland), Sweden, Switzerland (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Male.
Female.
Male.
Male.
Male.
Male.
Genus: Sintula Simon, 1884
Sintula retroversus (O. P.-Cambridge, 1875)
Range: Bulgaria, Croatia, France (Corsica), France (Mainland), Great Britain (Channel Islands), Greece (Dodecanese Islands), Greece (Mainland), Hungary, Italy (Mainland), Italy (Sardinia), Macedonia, Slovakia, Slovenia, Ukraine (van Helsdingen 2009.1). Global range: Europe to Azerbaijan (Platnick 10.0).
Genus: Styloctetor Simon, 1884
Styloctetor romanus (O. P.-Cambridge, 1872)
Range: Belgium, Bulgaria, Czech Republic, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Liechtenstein, Macedonia, Netherlands, Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Slovakia, Spain (Canary Islands), Spain (Mainland), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Male.
Male abdomen with spinners.
Male.
Male.
Female, epigyne.
Female.
Female.
Female.
Female.
Female.
Female.
Styloctetor stativus (Simon, 1881)
Range: Austria, Belarus, Belgium, Czech Republic, Denmark, Estonia, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Slovakia, Slovenia?, Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Holarctic (Platnick 10.0).
Genus: Tallusia Lehtinen & Saaristo, 1972
Tallusia pindos Thaler, 1997
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Genus: Tenuiphantes Saaristo & Tanasevitch, 1996
Biology: Previously, members of this genus were placed in the species rich genus of Lepthyphantes. A homogenous group of species have now been transferred to the Tenuiphantes genus which is characterised by their copulatory organs. Markings can be quite variable within species, nevertheless they are sometimes useful for separating the species.
Characters of genus: Small to large linyphiids ranging from 1.7-4.1 mm body length (Saaristo & Tanasevitch 1996). Males are characterized by having a sinuous embolus, often with a dentigerous protrusion at about halfway while females are characterized by having the proscapus bordered at either side by a lateral wing-like extension of the median part of the scapus (Saaristo & Tanasevitch 1996). Epigynes of T. alacris females also characterized by strongly developed lateral teeth. Paracymbium with 0-3 teeth. The species differ in leg spination. Metatarsus without a trichobothrium except in T. retezaticus (endemic to Romania). Most species have uniformly coloured legs and a dark dorsal pattern on the abdomen usually composed of broad black transverse bars on a brownish background. Bars sometimes reduced to paired dots combined by thin black lines. Dorsal pattern is usually more obscure in males. Legs fairly long and with long spines.
Tenuiphantes tenuis (Blackwall, 1852)
Description: Carapace brown to blackish. Anterior medials almost equidistant, with medials separated from laterals by ca. 0.5 diameter, laterals much less than twice the diameters of medials (Locket & Millidge 1953). Sternum blackish. Legs yellow-brown, fairly long and with long spines. TM 1 ca. 0.18-0.22 (Roberts 1987). Abdomen yellow-brown to almost black. Usually, dark transverse bars are present dorsally but they may be difficult to discern in specimens with dark background colours. Bars are often reduced to paired dots which may be combined by thin U- or V-bent black lines. Shining white patches are sometimes distributed across the dorsal surface of the abdomen, at other times mostly at sides if not absent completely. Epigyne anchor shaped, male palp with two teeth at each side of the paracymbium. Size: Female 2.0-3.2 mm; male 2.0-2.7 mm. Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Faroe Islands, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Greece (North Aegean Islands), Hungary, Ireland, Italy (Mainland), Italy (Sicily), Latvia, Liechtenstein, Macedonia, Malta, Moldova, Netherlands, Norway (Mainland), Poland, Portugal (Azores), Portugal (Madeira), Portugal (Mainland), Romania, Russia (Central European), Slovakia, Slovenia?, Spain (Balearic Islands), Spain (Canary Islands) (introduced), Spain (Mainland), Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Europe, North Africa (elsewhere, introduced) (Platnick 10.0).
Female.
Male.
Male.
Female.
Female.
Female.
Female.
Male.
Male.
Male.
Male.
Male.
Female.
Female.
Female.
Female.
Female abdominal markings.
Female, killed by parasitic larva shortly after final moult.
Male.
Male.
Male.
Male.
Male.
Male.
Genus: Tiso Simon, 1884
Characters of genus: Male head only slightly raised behind eyes. Eyes rather small. Metatarsus IV without a trichobothrium. Tm I 0.5-0.59 (Roberts 1987). Tibiae I-IV with one spine.
Tiso vagans (Blackwall, 1834)
Description: Carapace brown to blackish (generally darker in males which also have surface somewhat reticulated (not visible with a lens). Abdomen brownish-grey to black. Legs yellow-brown to greyish-brown. Tm I 0.51-0.59 (Roberts 1987). Male palpal tibia with some stout bristles and femur and patella are both elongated. Size: Female 1.7-2.2 mm; male 1.6-2.0 mm. Range: Andorra, Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Faroe Islands, Finland, France (Corsica), France (Mainland), Germany, Great Britain (Channel Islands), Great Britain (Mainland), Great Britain (Northern Ireland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (NW. European), Russia (Southern European), Slovakia, Slovenia, Spain (Mainland), Sweden, Switzerland, Yugoslavia (van Helsdingen 2009.1). Global range: Europe, Russia (Platnick 10.0).
Male.
Female.
Female.
Male.
Male.
Male.
Genus: Tmeticus Menge, 1868
Characters of genus: This genus description is based on Locket & Millidge (1953) and Saito & Ono (2001) and may not encompass all species currently assigned to this genus. Male head not elevated in to lobe but head may be highly raised. Male chelicerae with a large warty tooth anteriorly. Metatarsus IV with a trichobothrium. Female tibiae I-II with two spines, tibiae III-IV with one spine. Male tibiae I-II spineless, tibiae III-IV with one short spine. Male palpal tarsus and palpal organs are relatively small in some species. The species may be classified in to two groups based on the structure of the male palpal organs (Saito & Ono 2001). One group is characterized by very long male palps and a hook-shaped patellar apophysis while tibia is without distinct apophysis (except in Tmeticus affinis which have a small dorso-distal bifid tibial apophysis). The second group possess a shorter male palp without a patellar apophysis, but with a distinct dorsal apophysis on the tibia.
Tmeticus affinis (Blackwall, 1855)
Description: Thoracic part of carapace yellow-brown, cephalic part in front of cervical furrow blackish. However, according to Locket & Millidge (1953) the carapace is reddish-brown, rather darker in male. Locket & Millidge (1953) make no mention of a two-coloured carapace so perhaps uniformly coloured carapaces occur as well. Male chelicerae with a large warty tooth anteriorly and several small warts on front and sides, these are less pronounced in females. Sternum orange-yellow with many hairs. Legs yellow-brown, Tm I 0.65-0.75. Abdomen black with four distinct sigilla. Male palpal patella and tibia elongated while tarsus and palpal organs are relatively small . Size: Female 2.5-3.0 mm; male 2.5-2.7 mm. Range: Belgium, Czech Republic, Denmark, Estonia, Faroe Islands?, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Ireland, Latvia, Lithuania, Macedonia, Netherlands, Norway (Mainland), Poland, Russia (Central European), Russia (Northern European), Russia (NW. European), Slovakia, Sweden (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Genus: Trichoncus Simon, 1884
Trichoncus affinis Kulczynski, 1894
Range: Austria, Bulgaria, Croatia, Czech Republic, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Lithuania, Macedonia, Moldova, Netherlands, Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (NW. European), Slovakia, Slovenia, Spain (Mainland), Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Trichoncus hackmani Millidge, 1955
Range: Austria, Belgium, Bulgaria, Czech Republic, Denmark, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Italy (Mainland), Macedonia, Netherlands, Poland, Slovakia, Slovenia, Sweden (van Helsdingen 2009.1). Global range: Central, Northern Europe (Platnick 10.0).
Female.
Male.
Male.
Male.
Male.
Female.
Female.
Female.
Trichoncus vasconicus Denis, 1944
Range: Austria, Estonia, France (Mainland), Greece (Mainland), Norway (Mainland), Poland, Romania, Russia (Eastern European), Spain (Mainland), Ukraine (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Genus: Trichopterna Kulczynski, 1894
Trichopterna krueperi (Simon, 1884)
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Genus: Troxochrus Simon, 1884
Troxochrus scabriculus (Westring, 1851)
Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Netherlands, Norway (Mainland), Poland, Portugal (Mainland), Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Russia (Southern European), Slovakia, Sweden, Switzerland, Ukraine, Yugoslavia (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Female.
Female.
Genus: Typhochrestus Simon, 1884
Typhochrestus dubius Denis, 1949
Range: France (Mainland), Greece (Mainland) (van Helsdingen 2009.1). Global range: France (Platnick 10.0).
Genus: Walckenaeria Blackwall, 1833
Characters of genus: The members of this genus range in size from 1.35 to 4.0 mm (Millidge 1983). The male carapace in most European species is elevated, often into large lobes or modified in some other way. However, in a few species such as in W. dysderoides the male head is only slightly domed behind the eyes. Where there is a lobe this carries the posterior median eyes (Millidge 1983). In other species the male carapaces carries a projection, often furnished with hairs which may be clavate or furcated. More rarely is the carapace of the female elevated, such as in W. acuminata which carries a conical elevation. The sternum is longer than wide with the posterior end pointed between coxae IV (Locket & Millidge 1953). The pedicel is distinctly sclerotized and is quite conspicuous in some species. The abdomen is without a scutum and is unicoloured in most species, usually greyish black but occasionally light grey or yellowish brown. Tibia I and II carries two spines while III and IV carries one in the European species. Legs are unicoloured in most species, often bright orange or reddish orange. Some species have contrastingly blackened tibiae on anterior leg pairs. Spines are weak, particularly on legs I and II in males (Millidge 1983). All metatarsi with a trichobothrium, Tm I variable, ranging from 0.39-0.76 in British species (Roberts 1987). The male palpal organs are of similar form and differences are not discernible with a hand lens. Females of a few species possess characteristic epigynes which makes them identifiable in the field using a hand lens. Other characteristics of the genus (not visible with a lens) include the strongly pectinate and large superior tarsal claws of legs I and II, clear transverse striae on the lateral faces of the chelicerae and the acuminate tarsus of the female palp (Locket & Millidge 1953, Millidge 1983). The European members of the genus have been reviewed by Wunderlich (1972) and the North American by Millidge (1983).
Walckenaeria alticeps (Denis, 1952)
Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland), Hungary?, Italy (Mainland), Lithuania, Netherlands, Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Europe (Platnick 10.0).
Male.
Male.
Male.
Walckenaeria angustifrons (Simon, 1884)
Range: France (Mainland), Greece (Mainland)? (van Helsdingen 2009.1). Global range: France (Platnick 10.0).
Walckenaeria cavernicola Wunderlich, 1992
Range: Greece (Mainland), Spain (Canary Islands) (van Helsdingen 2009.1). Global range: Canary Is (Platnick 10.0).
Walckenaeria christae Wunderlich, 1995
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Greece (Platnick 10.0).
Walckenaeria claviloba Wunderlich, 1995
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: Crete (Platnick 10.0).
Walckenaeria dysderoides (Wider, 1834)
Range: Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France (Mainland), Germany, Great Britain (Mainland), Greece (Mainland)?, Hungary, Ireland, Italy (Mainland), Latvia, Liechtenstein, Lithuania, Netherlands, Norway (Mainland), Poland, Romania, Russia (Central European), Russia (Eastern European), Russia (Northern European), Russia (NW. European), Slovakia, Spain (Mainland), Sweden, Switzerland, Ukraine (van Helsdingen 2009.1). Global range: Palearctic (Platnick 10.0).
Female.
Female.
Female.
Female.
Male.
Male.
Walckenaeria exilis Blackwall, 1853 (Nomen dubium)
Range: Greece (Mainland) (van Helsdingen 2009.1). Global range: England (Platnick 10.0).